Status: Not recorded from Britain since 1973; widespread but very local in Ireland.
RDB Category: Endangered
Habitat: Wet humus in fens and bogs.
Other notes: Tarsal claw joints without ventral setae. Head with deep, curved frontal furrows anteriorly. Elytra parallel-sided, striae finely punctured, outer striae very faint; third interval with three or four deep fovate punctures. Wings present.

Status: Declining. Local and usually coastal in the extreme south and east of England.
RDB Category: Vulnerable
Habitat: Cracks and crevices, usually in clay soils or cliff bases but also on sandy or gravelly soils.
Other notes: Dorsal surface flat and pubescent throughout. Antennae entirely pubescent, basal segment wider than the remainder. Tarsal claws not toothed. Head sparsely but very strongly punctured; strongly contracted behind the protruding eyes, which have a strong row of curved hairs behind. Elytra densely punctured, intervals flat; apices transversely truncate. Wings present.